Peer-reviewed scientific publications
  • Van der Meij, J., Martinez-Gonzalez, D., Beckers, G.J.L.* and Rattenborg, N.C.* (2019) Intra-‘cortical’ activity during avian non-REM and REM sleep: variant and invariant traits between birds and mammals. SLEEP, 42:2. [doi:10.1093/sleep/zsy230] * co-senior and corresponding authors
  • Beckers, G.J.L., van der Meij, J, Lesku, J.A. and Rattenborg, N.C. (2014) Plumes of neuronal activity propagate in three dimensions through the nuclear avian brain. BMC Biology, 12:6. [doi:10.1186/1741-7007-12-16].
  • Beckers, G.J.L. and Gahr, M. (2012) Large-scale synchronized activity during vocal deviance detection in the zebra finch auditory forebrain. Journal of Neuroscience, 32(31): 10594-10608 [doi:10.1523/jneurosci.6045-11.2012].
  • Beckers, G.J.L., Nelson, B.S. & Suthers, R.A. (2004) Vocal tract filtering by lingual articulation in a parrot. Current Biology, 14, 1592-1597. [pdf] See also news articles on [Science], [Nature], and [New Scientist] websites or columns in [New York Times], [Geo] (in German) and [NRC] (in Dutch).
  • Beckers, G.J.L., Suthers, R.A. and ten Cate, C. (2003) Pure-tone birdsong by resonance filtering of harmonic overtones. Proceedings of the National Academy of Sciences USA, 100 (12): 7372-7376. [pdf] See also columns in [New York Times] and [NRC] (in Dutch).
  • Van der Meij, J., Ungurean, G., Rattenborg, N.C.*, and Beckers, G.J.L.* (2020) Divergent neuronal activity patterns in the avian hippocampus and nidopallium. European Journal of Neuroscience. [doi: 10.1111/ejn.14675]
  • Van der Meij, J., Rattenborg, N.C.*, and Beckers, G.J.L.* (2020) Evolution of sleep in relation to memory - A birds’ brain view. Current Opinion in Behavioral Sciences, 33, 78-85.[doi: 10.1016/j.cobeha.2019.12.004]
  • Kriengwatana, B.P. and Beckers, G.J.L. (2019) Speech perception: what do nonhuman animals have to say? In Human Language: From Genes and Brains to Behavior. Hagoort (ed.), MIT Press.
  • Rattenborg, N.C., van Der Meij, J., Beckers, G.J.L., and Lesku, J.A. (2019) Local aspects of avian non-REM and REM sleep. Frontiers in Neuroscience, section Sleep and Circadian Rhythms, 13, 567-- [doi:10.3389/fnins.2019.00567].
  • Trotter, A.S., Monaghan, P., Beckers, G.J.L., Christiansen, M.H. (in press, 2019) Exploring Variation Between Artificial Grammar Learning Experiments: Outlining a Meta-Analysis Approach. Topics in Cognitive Science Available at:
  • Van der Meij, J., Martinez-Gonzalez, D., Beckers, G.J.L.* and Rattenborg, N.C.* (2019) Neurophysiology of avian sleep: comparing natural sleep and isoflurane anesthesia. Frontiers in Neuroscience, Sleep and Circadian Rhythms, 13, 262-- [doi:10.3389/fnins.2019.00262 ] * co-senior and corresponding authors.
  • Tisdale, R.K., Lesku, J.A., Beckers, G.J.L.*, Rattenborg, N.C.* (2018) Bird-like propagating brain activity in anesthetized Nile crocodiles. SLEEP, 41(8), zsy105. * co-senior and corresponding authors [doi:10.1093/sleep.zsy105]
  • Tisdale, R.K., Tieri, L., Rattenborg, N.C., Beckers, G.J.L., and Lesku, J.A. (2018) Spectral properties of brain activity under two anesthetics and their potential for inducing natural sleep in birds. Frontiers in Neuroscience, Sleep and Circadian Rhythms, 12. [doi:10.3389/fnins.2018.00881]
  • Tisdale, R.K., Lesku, J.A., Beckers, G.J.L., Vyssotski, A.L, Rattenborg, N.C. (2018) The low-down on sleeping down low: Pigeons shift to lighter forms of sleep when sleeping near the ground. Journal of Experimental Biology, 221 (19). [doi:10.1242/jeb.182634]
  • Bolhuis, J.J., Beckers, G.J.L., Huybregts, M.A.C., Berwick, R.C., Everaert, M.B.H. (2018) The slings and arrows of comparative linguistics. PLoS Biology. [doi:10.1371/journal.pbio.3000019 ].
  • Bolhuis, J.J., Beckers, G.J.L., Huybregts, M.A.C., Berwick, R.C., Everaert, M.B.H. (2018) Meaningful syntactic structure in songbird vocalizations? PLoS Biology. [doi:10.1371/journal.pbio.2005157 ].
  • Beckers, G.J.L., Berwick B.C., Okanoya, K. and Bolhuis, J.J. (2017) What do animals learn in artificial grammar studies? Neuroscience & Biobehavioral Reviews [doi:10.1016/j.neubiorev.2016.12.021].
  • Beckers, G.J.L. and Rattenborg, N.C (2015) An in depth view of avian sleep. Neuroscience & Biobehavioral Reviews, 50:120-127. [doi:10.1016/j.neubiorev.2014.07.019].
  • Beckers, G.J.L., Berwick, B.C. and Bolhuis, J.J. (2014) Comparative analyses of speech and language converge on birds. Behavioral and Brain Sciences.
  • Berwick, B.C., Beckers, G.J.L., Okanoya, K. and Bolhuis, J.J. (2012) A bird’s eye view of human language evolution. Frontiers in Evolutionary Neuroscience, 4: 5 [doi:10.3389/fnevo.2012.00005].
  • Beckers, G.J.L., Bolhuis, J.J., Okanoya, K. and Berwick, R.C. (2012) Birdsong neurolinguistics: Songbird context-free grammar claim is premature. Neuroreport, 23(3): 139-145 [doi:10.1097/WNR.0b013e32834f1765].
  • Ohms, V.R., Beckers, G.J.L., ten Cate, C., and Suthers, R.A. (2012) Vocal tract articulation revisited: the case of the monk parakeet. Journal of Experimental Biology, 215: 85-92 [doi:10.1242/​jeb.064717]. See also news shot at [Science] website.
  • Berwick, R.C., Okanoya, K., Beckers, G.J.L. and Bolhuis, J.J. (2011) Songs to syntax: the linguistics of birdsong. Trends in Cognitive Sciences, 15(3): 113-121. [doi:10.1016/j.tics.2011.01.002].
  • Beckers, G.J.L (2011) Bird speech perception and vocal production: a comparison with humans. Human Biology, 83(2): 191-212.
  • Beckers, G.J.L. and Gahr, M. (2010) Neural processing of short-term recurrence in songbird vocal communication. PLoS ONE, 5(6): e11129 [doi:10.1371/journal.pone.0011129].
  • Ohms, V.R., Snelderwaard, P.C., ten Cate, C. and Beckers, G.J.L. (2010) Vocal tract articulation in zebra finches. PLoS ONE, 5(7): e11923 [doi:10.1371/journal.pone.0011923].
  • Beckers, G.J.L. (accepted) Peripheral mechanisms of vocalization in birds: a comparison with human speech. In: Birdsong, Speech and Language. Converging mechanisms, Bolhuis, J.J. and Everaert, M. (eds.). MIT press.
  • Ohms, V.R., Gill, A., Van Heijningen, C.A.A. Beckers, G.J.L. and ten Cate, C. (2009) Zebra finches exhibit speaker-independent phonetic perception of human speech. Proc. R. Soc. B [doi: 10.1098/rspb.2009.1788] See also research highlight in [Nature], and news focus at [Science] website.
  • Beckers, G.J.L. and ten Cate, C. (2006) Nonlinear phenomena and song evolution in Streptopelia doves. Acta Zoologica Sinica, 52: 482-485. [pdf]
  • Beckers, G.J.L. (2006) Vocal production in birds. In The Encyclopedia of Language and Linguistics., 2nd edition, Brown, K. (Ed.), pp. 454-457, Oxford: Elsevier.
  • Suthers, R.A., Beckers, G.J.L. & Nelson, B.S. (2005) Vocal Mechanisms for Avian Communication. In Behavior and Neurodynamics for Auditory Communication Kanwal, J.S. & Ehret, G. (Eds.) Cambridge: Cambridge University Press.
  • Nelson, B.S., Beckers, G.J.L. & Suthers, R.A. (2005) Vocal tract filtering and sound radiation in a songbird. Journal of Experimental Biology, 208 (2): 297-308. [pdf]
  • Fletcher, N.H., Riede, T., Beckers, G.J.L. & Suthers, R.A. (2004) Vocal tract filtering and dove coos. Journal of the Acoustical Society of America, 116(6), 3237-3797. [pdf]
  • Riede, T. Beckers, G.J.L., Blevins, W. & Suthers, R.A. (2004) Inflation of the esophagus and vocal tract filtering in ring doves. Journal of Experimental Biology, 207 (23): 4025-4036. [pdf]
  • Beckers, G.J.L., Suthers, R.A. & ten Cate, C. (2003) Mechanisms of frequency and amplitude modulation in ring dove song. Journal of Experimental Biology: 206, 1833-1843. [pdf]
  • Beckers, G.J.L., Goossens, B.M.A., and ten Cate, C. (2003) Perceptual salience of acoustic differences between conspecific and allospecific vocalizations in African collared-doves. Animal Behaviour, 65: 605-614. [pdf]
  • Beckers, G.J.L. & ten Cate, C. (2001). Perceptual relevance of species-specific differences in acoustic signal structure in Streptopelia doves. Animal Behaviour: 62, 511-518. [pdf]
  • Verhoeven, K.J.F. and Beckers, G.J.L. (1999) Canopy farming: an innovative strategy for the sustainable use of rain forests. Selbyana, 20 (1): 191-193.
  • Beckers G.J.L., Leenders T.A.A.M. & Strijbosch H. (1996). Coral snake mimicry: Live snakes not avoided by a mammalian predator. Oecologia, 106 (4): 461-463.
  • Leenders, T.A.A.M., Beckers, G.J.L. & Strijbosch, H. (1996). Micrurus mipartitus (NCN) polymorphism. Herpetological Review 27(1): 25.
Book reviews
  • Beckers, G.J.L. (2009) Neuroscience of Birdsong. Edited by H. Philip Zeigler & Peter Marler. Cambridge: Cambridge University Press, 2008. Animal Behaviour [doi:10.1016/j.anbehav.2009.03.009].
  • Beckers, G.J.L. (2009) Birdsong: Chapter and Second Verse. Review of: Bird Song: Biological Themes and Variations. Second Edition. C.K. Catchpole & P.J.B. Slater Cambridge: Cambridge University Press, 2008. Ethology, 115: 511-512.
  • Beckers, G.J.L. (2006) Review of: The Physics of Birdsong. G.B. Mindlin & R. Laje. Berlin: Springer-Verlag, 2005. Animal Behaviour, 71: 995-996. [pdf]
  • Beckers, G.J.L. & Bolhuis, J.J. (2000) Review of: The Design of Animal Communication. M.D. Hauser & M. Konishi (eds.). Cambridge, MA: MIT Press, 1999. Animal Behaviour, 60: 703-704. [pdf]
  • Beckers, G.J.L. (2003) Vocal production and perception in Streptopelia doves. PhD thesis Leiden University, isbn 90-6734-036-7. [pdf]